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Funding information National Institutes of Health, Grant/Award Numbers: 5T32AG021885, NS043277 Abbreviations: ALS amyotrophic lateral sclerosis AMPA alpha-amino-3-hydroxy-5-methyl-4-isoxazolepropionic acid BCS bathocuproine disulfate CCS copper chaperone for superoxide dismutase COG complex conserved oligomeric Golgi complex CPZ cuprizone Cu (II)-atsm diacetylbis(4-methylthiosemicarbazonato)copper (II) CuES copper elescomol CuHIS copper histidine DBH dopamine β -hydroxylase dHMN X-linked distal hereditary neuropathy DTT dithiothreitol EAE experimental autoimmune encephalomyelitis GSH glutathione GSSG glutathione disulfide LC locus coeruleus MS multiple sclerosis NE norepinephrine NMDA N-methyl-d-aspartate OHS occipital horn syndrome PAM peptidylglycine α -amidating PrP C cellular prion protein RA retinoic acid REM rapid eye movement SCN suprachiasmatic nucleus SOD1 superoxide dismutase 1 Steap six-transmembrane epithelial antigen of the prostate TGN trans -Golgi network TTM tetrathiomolybdate WD Wilson disease Footnotes CONFLICT OF INTEREST STATEMENT The authors declare no competing financial interests